1940年3月，汪伪政府宣布成立。侵华日军在上海大肆搜捕抗日志士，汪伪组建由前国民党军官毕忠良担任处长的“特别行动处”为虎作伥。一直潜伏在毕忠良身边、代号“麻雀”的中共党员陈深，与组织暂时失去联系。
置身虎视狼顾的险境中，陈深信仰坚定，冒死营救上线“宰相”及抗日志士。毕忠良老奸巨猾，虽与救命恩人陈深“情同兄弟”，却对其产生怀疑，设重重陷阱步步紧逼。“宰相”牺牲后，陈深终于找到新上线“医生”并受命拿到日军代号为“归零”的重要作战计划。
陈深的昔日恋人徐碧城，与另一国军情报人员唐山海假扮夫妻潜伏进特别行动处，在多次行动中给陈深制造麻烦和危机。智计过人的陈深身处连环杀机之中，巧妙利用日本鬼子、汪伪分子等各派势力之间的微妙缝隙，同时也付出了巨大的代价，最终成功地获取了“归零”计划，挫败了日本侵略者的阴谋。
麻雀第69集
毕太太的生日宴会热闹非凡，行动处队员们大肆喝起酒来，而刘二宝却紧紧盯着兄弟们的一举一动，扁头也没有喝酒，他知道陈深今晚一定有所行动。此时，徐碧城正准备趁机离开上海，她拎着箱子快步走在街上，却被突然出现的陶大春拦住了去路。陶大春不能忍受徐碧城被共党策反，更不能允许她和陈深在一起，于是将碧城打晕，企图阻…
麻雀第68集
苏三省受伤后慌忙逃跑，躲进村郊一户农家中，看着自己的伤口，苏三省发誓一定不会放过毕忠良和陈深。李默群向影佐建议立即将苏三省捉拿归案，这一次，李默群确实大力支持了毕忠良，虽然他并不理解毕忠良要杀苏三省的意图。可以确定的是，苏三省抓捕唐山海和徐碧城功，重重打击到了李默群，即使没有毕忠良，李默群也是想将苏…
麻雀第67集
毕忠良将“回家计划”拿给陈深看，旁敲侧击地询问皮皮是否与这个计划有关。陈深离开行动处后，心中忐忑不已，他不知道这是否是毕忠良设下的圈套，但无论如何，他还是决定去审问庞勇，不放过任何拯救皮皮的机会。毕忠良知道皮皮现在在苏三省手中，为了逼陈深暴露，决定再次与苏三省联手。苏三省对陈深恨之入骨，当然愿意与毕…
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