白天里，热热闹闹的动物园里，所有的动物伙伴们都使劲浑身解术，弄得前来观看玩耍的大朋友和小朋友们不亦乐乎。而当夜幕降临的时候，动物们却依然有着足够的精神头，不会虚度一点光阴，等到动物园打烊之后，来自地球各处的各种动物们又开始了自己的狂欢派对，一同尽情地享受着玩闹、嬉闹的乐趣，那场面更胜过人类所有的狂欢节日让人忍俊不禁。但是，突如其来的事件，打乱了动物们原本欢乐幸福的生活。可爱调皮的小狮子瑞恩在与日俱增的好奇心的驱使下，勇敢地爬上了一辆远去的货车车箱，并随着货车上的货物要一同被辗转带到遥远的非洲去。这可急坏了狮子爸爸，为了救回心爱的儿子瑞恩。狮子爸爸号召了众多动物园里的好朋友们前来帮忙。长颈鹿、考拉、水蟒和松鼠都积极参加了这个由狮子爸爸领头的这支特殊的“寻子远征队”。经过一番准备之后，远征队雄纠纠地踏上了长途寻子的旅程。这群在动物园中养尊处优的动物远征队上天入地，要找回自己的伙伴，自然不会一帆风顺，途中遇到了各种各样的危险：在大街上被野狗追咬；在空中坐着海鸥翱翔；在海上乘船差点被撞沉；还莫名其妙的进了下水道，几乎丧命于鳄鱼之口，真是艰险重重，再加上几个伙伴个个都不是省油的灯，在漫长而有趣的旅途中上演了一幕幕有惊无险的闹剧……
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