北约特工雅克·克里斯托夫（Jacques Kristoff）在他的生日被召唤起来，追捕加林娜·康斯坦丁（Galina Konstantin），后者以极其珍贵和危险的绝密货物为生。找到Galina并不需要很长时间，Jacques必须通过乘火车将Galina和违禁品交还给上司来完成任务。雅克的内科医生妻子玛德琳（Madeline）对这种情况不满意。但她却看到他和这对夫妇的十几岁的女儿贝利·克里斯托夫（Bailey Kristoff）和儿子埃森·克里斯托夫（Ethan Kristoff）在火车站下车。 Galina的不义之财是三瓶SP-43，这是一种超强毒天花菌株，在含有其他几种病原体的荧光绿色血清中培养。由梅森·科尔（Mason Cole）领导的一群恐怖分子打算劫持火车，并为了自己的目的窃取病毒。但是雅克挡住了他们的路。病毒最终在火车的通风系统中散失。不幸的是，雅克的家人选择了这一刻，以暗中加油骑行为他的生日感到惊讶。雅克为了保护家人和其他乘客并阻止科尔和他的追随者而成为一支单人军队
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