东京的平凡角落，普普通通的佐藤家兄妹相守度日。初中毕业干起修路工大哥佐藤旭（妻夫木聪饰）凭一己之力拉扯弟妹长大，因共同的爱好他和美丽善良的盒饭店服务员泽边梓（苍井优饰）走到一起。当年因诈骗入狱的老二晓（瑛太饰）重获自由，几经辗转在受害者的女儿屋代多香子（长泽雅美饰）的田里帮忙。唯一一个女孩阿光（满岛光饰）担任妇产科护士，与和大哥从小一起长大的医生新城正臣（吉冈秀隆饰）发展出不伦关系。热爱表演不惜放弃学业的佐藤阳（柄本佑饰）和弟弟旦（野村周平饰）同在一个学习班的女孩永原香澄（桥本爱饰）偶然相逢，他将这个擅长表演的女孩拉入自己的剧团。佐藤家的孩子执拗善良，冲动率真，兄妹们用各自的人生为新世纪的年轻人们写下注脚……
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