穿过如絮的云层俯瞰大地，田畴如茵阡陌纵横。在这绿草如茵的地底下，胆小的鼹鼠在他狭小温馨的家里正筹备过冬的物资。让鼹鼠始料不及的是，一场突如其来的灾难正在他的头顶向他袭来。隆隆的铲车碾过后，还没等鼹鼠搞清楚发生了什么，鼹鼠温馨的小家顷刻间被压塌。
无家可归的鼹鼠在河边遇到了河鼠。听说了鼹鼠的不幸遭遇，河鼠热情地挽留鼹鼠在自家居住，两人同行结伴，沿着河流旅游。河鼠的盛情挽留被鼹鼠拒绝，鼹鼠想知道究竟是谁毁了他温馨的家。得知家上面那块草坪是蟾蜍的，家很有可能是蟾蜍毁坏的后，鼹鼠在河鼠的陪伴下，来蟾宫找蟾蜍问个究竟。
蟾蜍凭借父辈创造的丰厚家业日子过得悠闲而富庶。尽管蟾蜍有自吹自擂的恶习和酷爱汽车的嗜好，但蟾蜍表明鼹鼠的家绝对不是他毁的。在蟾蜍驾车带鼹鼠和河鼠去旅游的路上，河鼠得知蟾蜍因手头紧张，早已经把那块草坪卖给了黄鼠狼。蟾蜍追求极速的快感给他和他的朋友...
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